Agreed. Feet look clean and healthy, and if they were removed from shoes, I can guarantee they have less foreign matter on them then the shoes they were removed from, so I’d prefer them on the seat cushion rather than the shoes. If they were removed from shoes, they might be sweater and have more bodily products in general than the shoes they were removed from, but I doubt the seats on public transit are going to long avoid sweat and skin.

Some people’s insistence on covering all body parts at all times in public makes no sense to me.